Existing and emerging uses of drones in restoration ecology

In the absence of effective and scalable human intervention, up to 95% of the world's ecosystems will be affected by anthropogenic degradation by 2050. Therefore, immediate and large‐scale ecological restoration is imperative to stem biodiversity loss and ecosystem decline. Ecologists must draw upon the most effective and efficient tools available to achieve successful restoration goals. Drones (i.e., unmanned aerial vehicles) are a valuable set of tools in the environmental, forestry, and agriculture sectors; however, there has been limited uptake in restoration ecology. Here, we aim to highlight the existing and emerging uses of drones in restoration science and practice. We discuss the strengths and weaknesses of these applications and provide a roadmap for increasing the utilisation of drones to refine and enhance restoration objectives. Our article is presented with the restoration continuum in mind, including sections for restoration planning, implementation and monitoring. We also take a novel approach by describing how drones relate to a globally recognised restoration tool published by the Society for Ecological Restoration. Drones are used in several restoration scenarios from mapping habitats and managing wildfires, to monitoring the effectiveness of restoration interventions. Many applications in other disciplines can also be transferred to restoration scenarios. However, the use of drones will be context‐dependent, and several technical and practical constraints need to be addressed. Drones have considerable potential to improve the science and practice of restoration at all stages of a restoration project, which is vital to realising the goals of the UN Decade on Ecosystem Restoration.

Drones (also called unmanned aerial vehicles [UAVs]) have rapidly advanced over the last decade and are now primed to help tackle the complexities of ecological restoration. There are two main types of drones: (a) fixed-wing and (b) multi-rotor (Box 1). A diverse range of sensor units are now available for fully integrated and tailored drones, which can combine drones with artificial intelligence and provide greater autonomy, precision and efficiency of practice. Available sensors include active sensors, such as light detection and ranging (LiDAR) modules, which use laser-based technology to generate very high-density point clouds (e.g., >1,000 points/m 2 ) to detect objects at fine scales such as individual trees to model provenance performance differences (Camarretta et al., 2020). Passive sensors are also available, such as optical sensors, including high-definition cameras that can be used in combination with object recognition algorithms to monitor species and ecological communities (Dalla Corte et al., 2020;Hamylton et al., 2020;Harrison, Camarretta, et al., 2021;Lu et al., 2020). In addition, drones can carry other payloads that are not sensors. These can include units that disperse fire extinguishing materials to help control wildfires, fire starting materials for prescribed burns, or specialised seed packages apparatus designed to disperse seeds and plant amendments (e.g., fertilisers) to meet revegetation objectives (Aydin et al., 2019;Droneseed, 2020;Rahman et al., 2019;Saikin et al., 2020).
However, compared to these disciplines, the uptake of drones by the restoration sector has been limited, despite their increasing versatility and affordability (Buters, Bateman, et al., 2019;Camarretta et al., 2020). This lack of uptake could be attributed to several factors, including methodological ambiguity, a lack of standardised guidelines, and variations in policy frameworks--which may also compound the effects of more general concerns, such as perceptions around safety and intrusiveness (Buters, Bateman, et al., 2019;Rao et al., 2016).
To maximise the benefits of drones in restoration, it is important that end-users are aware of their strengths and weaknesses and the need for further innovation and development. To this end, we aim to (a) understand the extent to which drones are currently used in restoration and highlight emerging applications, (b) identify opportunities to transfer drone-based approaches from other disciplines (e.g., agriculture, conservation biology) into restoration, (c) examine key barriers to their broader adoption and (d) discuss how drones fit into the Society for Ecological Restoration's (SER) international principles and standards for the practice of ecological restoration (Gann et al., 2019). SER 'advances the science, practice, and policy of ecological restoration to sustain biodiversity, improve resilience in a changing climate, and reestablish an ecologically healthy relationship between nature and culture' (SER, 2022). Their standards guide restoration practitioners and help ensure that time and resources invested in restoration projects are well spent. We provide a restoration-centric narrative, focusing on the existing and potential uses of drones at three key stages of a restoration project: planning, implementation and monitoring (Figure 1), whilst anchoring to the SER ecological restoration 'recovery wheel'.

| THE S ER RE S TOR ATI ON RECOVERY WHEEL
The second edition of the international principles and standards for the practice of ecological restoration was released in 2019 (Gann et al., 2019). This publication guides the practice of ecological restoration and contains an ecological restoration 'recovery wheel' for assessing the success of a restoration project. The wheel measures the recovery of six distinct but interacting factors of an ecosystemspecies composition, structural diversity, ecosystem function, external exchanges, absence of threats and physical conditions. Each of these components are scored between 1 and 5, where 1 indicates the lowest level of recovery outcome, and five the highest. While the recovery wheel provides a mechanism to track the progression of an ecosystem's recovery towards a desirable condition and the initiation of adaptive management options to re-set the trajectory, many of the factors that are assessed can be costly and time consuming to gather, especially as restoration is upscaled from local to landscape-scales. However, the rapid evolution of drone technology is now making it possible to assess these ecosystem factors at the required scale, and we highlight multiple examples of how drones could be used to support each of the six ecosystem recovery factor ( Figure 2) through extensions of their applications in planning, implementing and monitoring restoration outcomes.

| RE S TOR ATION PL ANNING
Restoration projects require adequate planning to be successful.  (Gann et al., 2019;Harrison, Camarretta, et al., 2021;. Drones can have a demonstrable role in each step, which we outline below.

| Image-based vegetation mapping
Producing maps to acquire information on baseline ecological conditions is fundamental to planning subsequent restoration strategies (European Environment Agency, 2016). However, this can be timeconsuming and labour-intensive if traditional, on-ground surveys are undertaken. While satellite imagery can be used, it can have reduced temporal and spatial accuracy, and the resolution is sometimes not at the fine-scale required for restoration planning, however, dronesatellite synergies are being explored (Alvarez-Vanhard et al., 2021).
The potential for other remote-sensing applications in planning restoration approaches are emerging, such as capturing drone-derived imagery that produce large-scale topographic and vegetation maps through photogrammetry approaches (Cruzan et al., 2016). Careful checking for sources of noise that could confound the classification of target units for restoration is essential. In the case of the above example, validating the classification was required by Sierra-Escrigas et al. (2020) as initial variability was detected due to signal similarity and F I G U R E 1 Examples of applications for drones along the restoration continuum from planning and implementation to monitoring and adaptive management. (a) Indicates a fixed-wing drone system able to collect baseline ecological and geophysical conditions over large spatial scales; (b) shows a multirotor drone able to deploy seed packages and fertilisers; and (c) indicates a multirotor drone able to conduct fine-scale assessments of vegetation complexity (e.g., via canopy height models, species counts).
interfering components of the atmosphere and water column. Although this study was carried out in a conservation context, the principles are transferable to restoration ecology for mapping baseline conditions and delineating ecological units. The application of mapping units of interest extends to the terrestrial landscape where drones have been used to detect plant species to delineate vegetation units (Baena et al., 2017;Villoslada et al., 2020). There is a clear, demonstrated approach on the utility of drones to assist in identifying invasive species and key ecological species that has extended applications in a restoration planning context. For instance, controlling invasive species is often vital to ecosystem restoration. Decisions about which control methods to use depend on the plant species' or growth forms and their spatial distributions. Drones could be used to map these taxonomic, structural and spatial elements and therefore aid in restoration decision-making.
This contributes to several SER Recovery Wheel objectives (Figure 2), including providing a rapid assessment of the presence or absence of threats (Absence of Threats), or the baseline species composition and structural diversity of the ecosystem being restored.
Other studies have used off-the-shelf drones (e.g., DJI Phantoms) along transects to map intertidal, coastal (mangrove) and inland vegetation (Cruzan et al., 2016;Darmawan et al., 2020;Rossiter et al., 2020). Aerial images are collected by the drone and used to create orthomosaics and digital surface models (DSM) to map vegetation and landform elevation differences. The resulting orthomosaic can then be analysed using software such as ESRI ArcGIS 10.8 (ESRI, 2021), which has relevant processing tools and an 'Ortho Mapping' workspace. Similarly to the coral reef study mentioned above, ecological features displaying strong spectral or elevational differences allow for more accurate automated habitat delineation using DSMs.
These studies suggest that using drone-derived imagery can be valuable in a restoration context to, for example, acquire baseline ecological condition data and determine landscape prioritisation.
Future research to improve the accuracy of imagery-based vegetation maps is needed. In particular, there is a need for methodological guidelines that restoration ecologists using drones can apply across F I G U R E 2 The SER recovery wheel and some examples of how drones could potentially contribute to assessing the recovery of each ecosystem factor. For instance, in the Species Composition segment, drones could be used to plant and monitor the development of plant composition and diversity against a goal.For the Structural Diversity segment, drones could be used to understand vegetation strata and spatial distribution by facilitating digital models. The figure provides examples of how drones can be used in each of the other segments of the recovery wheel. The 5-star scale represents a cumulative gradient from very low to very high similarity to a reference ecosystem. A restoration site can be assigned to one of the five recovery levels (1-5 stars) as indicated by the green subsegments. different ecosystems that detail the optimal approaches and associated pitfalls.

| LiDAR data fusion for rapid phenotyping
Drones can now support LiDAR sensors to measure the structural properties of an ecosystem (e.g., forests) at high spatiotemporal resolution (Camarretta et al., 2020;Jaakkola et al., 2010;Wallace et al., 2014Wallace et al., , 2016 ing. The authors used the hierarchical density-based spatial clustering of application of noise clustering algorithm (Ester et al., 1996) to segment tree stems. Following this, the authors used a principal component analysis to extract tree stem orientation for subsequent diameter at breast height estimation. This workflow was validated using LiDAR point clouds collected (using a maximum scanning angle Range of 75°) in a temperate deciduous closed-canopy forest stand during the leaf-on and leaf-off seasons. The results suggested that this approach detected ~82% of tree stems (with a precision of 98%) during the leaf-off season. The authors state that the point density within an approximately 1.3-m height above the ground was low within closed-canopy forest stands, thereby reducing the accuracy of diameter at breast height estimation. In summary, a LiDAR-based detection level of 82% for tree stems during the leaf-off season suggests improvements to the accuracy of LiDAR-based tree mapping are still needed. Yet, data fusion approaches that combine hyperspectral and LiDAR data seem to be the most effective approach to assess forest structural attributes and tree species richness for restoration planning in a forest environment. Additional research on the use of drone-based LiDAR platforms to gain baseline information for non-forest ecosystems, such as grasslands, is needed to ascertain the benefits of data fusion to remotely detect and classify vegetation composition in various restoration contexts, thereby aiding in restoration planning.

| Baseline plant stress/health assessments
Drones mounted with multispectral cameras can be used to acquire data on plant health via vegetation indices such as the Normalised Difference Vegetation Index (NDVI) and the Enhanced Vegetation Index (EVI) (Figure 3). A considerable amount of work has been done on this in the agricultural sector. For example, Kim et al. (2021) used the drone NDVI approach to assess crop damage after chemical exposure. NDVI is commonly used to detect changes in plant health derived from the difference between visible and near-infrared reflectance of vegetation cover. Rice crops were exposed to chemicals at five growth stages to four levels of the chemical toluene. The NDVI was measured 5 days after damage and 67 days after planting. NDVI of toluene-exposed rice was significantly lower at most growth stages. The authors suggested that this approach indicated that NDVI assessed at close range, as enabled by the drone, could detect plant responses to toluene exposure.
Drone-based vegetation assessments that exploit indices such as the NDVI or Photochemical Reflectance Index (PRI, de- A key pitfall to only using drones is the reduced spatial coverage compared to satellites. This is something to consider in a restoration context. For example, does the restoration project require higher resolution/targeted indices or broader spatial coverage? There may also be challenges for time series analyses, whereby data points are recorded at consistent intervals over a set period of time. Satellites do this consistently, whereas additional project planning and management is required for frequency data collection using drones.

| Counting wildlife
A considerable body of research has demonstrated how drones can be effectively deployed to remotely count wildlife. These data can be used to establish baseline conditions (e.g., determining species presence and abundance) for restoration projects. Francis et al. (2020) developed a semi-automated wildlife counting method, using drones and machine learning to identify and count waterbird species in the Okavango Delta, Botswana and the Lowbidgee floodplain, Australia.
Their detection accuracy between the training and test data was 91% for the Okavango Delta colony and 98% for the Lowbidgee floodplain colony. In addition, their semi-automated method was 26% quicker, including development, and five-fold quicker without development than manual counting. This, as the authors emphasised, suggests that drone data of waterbird colonies can be collected rapidly and accurately, allowing for counting with minimal disturbance. This is corroborated by Davis (2021), who used a drone equipped with a thermal camera to detect deer in a range of habitats and to generate deer density estimates. The results showed that the drone outperformed ground observations regarding density, surveying efficiency, and area covered. Drones with thermal cameras are currently being used by the Woodland Trust in the UK for herbivore impact assessments. These assessments are vital in the planning stage of restoration projects. The drones allow forest restoration ecologists to efficiently count herbivore populations in the local area, and thus allow them to gauge herbivore pressure. This information is then used to formulate a tree protection plan, for example, do the saplings require tree guards, or is fencing enough? Drones can therefore contribute to several SER Recovery Wheel objectives

| RE S TOR ATION IMPLEMENTATION
The choice of restoration intervention will be context-dependent and generally should align with natural ecological processes, not cause any further degradation, and be compliant with local regulations (Gann et al., 2019). Here, we discuss how drones can and are used to implement restoration interventions.

F I G U R E 3
An example of how drones can monitor plant health is by utilising a modified multispectral camera to obtain normalised difference vegetation index (NDVI) values. Low NDVI values correspond to no or highly stressed vegetation, and high NDVI values correspond to higher levels of greenness as a proxy for relative chlorophyll output and therefore a proxy for high plant health. Panel (a) shows an orthomosaic image of a woodland-heathland ecotone in the Peak District, UK, and (b) visual representation of the NDVI-Where red indicates soil, rock and dead vegetation (Robinson, unpublished data using Pix4DMapper™, 2018).

| Seed dispersal and pest management
Despite very few peer-reviewed studies on drone-based seed dispersal, there is great enthusiasm for its potential to assist in revegetation projects (Mohan et al., 2021). Dispersing seeds (along with fertilisers) using drones has been applied to help revegetate mangroves in Myanmar and the United Arab Emirates and has assisted in revegetation operations in Thailand, where the landscape is inaccessible or unsafe for humans (Mohan et al., 2021). However, a recent study demonstrated that the survival percentage of seeds dispersed by drone was low and ranged from 0% and 20% for certain conifer species (Aghai & Manteuffel-Ross, 2020). In this pilot study, numerous environmental factors (e.g., humidity, solar exposure and predation) limited the establishment of the seedlings. Still, this low survival level is comparable to some traditional seed-based restoration plantings (e.g., <25% survival; Woods et al., 2019), whilst others are considerably higher such as an 83% success rate for seedling establishment in a recent dryland restoration project (Shackelford et al., 2021). Given the paucity of peer-reviewed operational studies, it is difficult to compare the effectiveness and potential of drone-assisted seed dispersal to traditional seeding approaches.
Nonetheless, drone-assisted seed dispersal does appear to have the potential to complement traditional planting efforts, particularly in dangerous or complex landscapes.
As pest species contribute heavily to biodiversity loss and degrade ecosystems, pest management is considered a key part of  (Li, Giles, Niederholzer, et al., 2021) and cranberry farms (Luck et al., 2021). It could be valuable to use drones in this way to reduce the threats associated with seed predation-which is a considerable issue for drone-supported seed sowing activities.

| Wildfire tracking and control
Although wildfires managed for restoration can positively impact ecosystem recovery (Barros et al., 2018;White & Long, 2019), unprescribed and/or uncontrollable wildfires can have a detrimental ecological impact (Lewis, 2020). Therefore, innovative methods to help detect, track and extinguish certain types of wildfires are being developed. Further research into drone-assisted prescribed burning would also be beneficial (Beachly et al., 2016).
Detecting and tracking wildfires with drones via thermal detection (e.g., using infrared cameras) has considerable potential (Allison et al., 2016;Athanasis et al., 2018;Kumar et al., 2011;Pham et al., 2018) and therefore potential to contribute to the SER Recovery Wheel objective Absence of Threats (Figure 2 et al. (2019) show that smaller sized fire extinguishing balls might be effective in extinguishing short grass fires (i.e., a ~0.5 kg ball effectively extinguished a circle of 1-m of short grass). This fire extinguishing potential of ball dispersing drones has been well-recognised, and many systems are currently being developed and tested (Bailon-Ruiz & Lacroix, 2020; Barua et al., 2020;Innocente & Grasso, 2019). We will likely have a much clearer picture of the effectiveness of drones to help extinguish wildfires in the coming years.

| RE S TOR ATION MONITORING
Revegetation plantings must be monitored to evaluate whether the project's goals are being met and inform adaptive management strategies to potentially re-set the trajectory of projects towards the desired goals (Gann et al., 2019). Monitoring a restoration project involves evaluating outcomes, which typically includes the status of the intervention (e.g., plant health, community complexity) and the degrading processes or ecological threats (e.g., pest populations).
Here, we discuss how drones can and are used to monitor restoration projects. Many drone applications and techniques used in the restoration planning and implementation stages (discussed above) can apply to restoration monitoring, and we only provide a relatively brief summary accordingly.

| Monitoring plant stress/health
Using the same methods to assess baseline plant health conditions

| Wildlife surveillance
Wildlife surveillance may be required as part of a restoration assessment-for example, to understand the wildlife responses to landscape repair (Jones & Davidson, 2016). Corcoran et al. (2021) found that automated wildlife detection using drones can be achieved for a wide range of species (e.g., koala Phascolarctos cinereus; glossy ibis Plegadis falcinellis) and under various ecological conditions (e.g., low dense eucalyptus forest, coastal wetlands). This is corroborated by several studies from different ecosystems, for example, coastal (Oosthuizen et al., 2020), marine (Dickens et al., 2021) and inland (Eori et al., 2020;Hui et al., 2021). Hollings et al. (2018 pointed out a potential constraint to using drones for automated wildlife detection is that accuracy often reduces over larger spatial scales, and the cost of high-resolution data is relatively high. Nonetheless, it has been estimated that using drones to survey 100 ha of land is ~10 times more efficient than surveys based on traditional field surveys (Filipovs et al., 2021). In addition, a recent study linked drone photogrammetry derived vegetation structural traits with animal tracking to understand the structural components required to elucidate animal behaviour (Harrison, Camarretta, et al., 2021;. Lee et al. (2021) pointed out that other approaches may be more efficient because automated detection systems require significant amounts of time to generate training data and train detection models. The authors proposed a real-time animal detection method based on the Sobel edge algorithm, which can detect animals in a single image without training data. In their feasibility study, the fastest detection time per image was 0.033 s. Target images were acquired at heights <100 m, and the maximum detection precision was 0.804. These studies highlight several ways of acquiring wildlife detection and counts that could be useful in a restoration context-that is, to understand the components of the ecosystem that has undergone a restoration intervention. Streamlining the analytical pipeline to make it easier for all operators (including non-specialists) to conduct automatic wildlife detection and interpretation will bring value to restoration ecology. Detecting wildlife using drones contributes to several SER Recovery Wheel objectives (Figure 2), including Absence of Threats, Species Composition, External Exchanges (e.g., gene flows) and Ecosystem Function (e.g., interactions and productivity).

| Bulk environmental sampling
Environmental sampling, such as collecting soil or water samples for downstream metagenomic or physicochemical analysis, can provide important information for restoration monitoring (Breed et al., 2019).
A multirotor drone mounted with landing gear and a sampling device could be a valuable tool in this regard. Collecting water samples to acquire environmental DNA (eDNA), preceding polymerase chain reaction (PCR) and analysis, can provide detailed data on ecological community composition and target species presence (e.g., pest or conservation priority species) in a waterbody (Rees et al., 2014).
Restoring an ecosystem might involve creating ponds to support aquatic wildlife. By using a drone-based water sampling system, eDNA samples can be collected from a given pond (particularly useful when the banks are steep or where marginal vegetation is dense, thus preventing on-foot sampling) to detect amphibians ( Figure 5).
This approach can help restoration practitioners to assess whether the pond restoration intervention has had the desired effect (e.g., Figure 2). Invertebrate diversity surveys can also be conducted in this manner (Nguyen et al., 2020).

Ecosystem Function and Species Composition;
Similar sample collection devices can be attached to drones to potentially acquire other types of environmental samples in terrestrial habitats-e.g., soil and invertebrates. For example, in dangerous or relatively inaccessible landscapes, drones could be used to collect soil to assess the microbial composition and physicochemical properties, thus allowing soil health and biotic communities to be monitored following a restoration intervention. For example, the NIMBUS Lab at the University of Nebraska developed a drone soil drilling system that could be adapted to collect soil cores for this purpose. Ecological consultants have used drones with a tick (parasitic invertebrate) flagging systems attached to survey Ixodid ticks in a meadow habitat (unpublished). This enabled the operator to fly a drone across a grassland to survey ticks safely (i.e., removing the surveyor from vector hotspots) and efficiently as part of an invertebrate DNA surveying project to detect elusive species such as the European hedgehog Erinaceus europaeus. Notably, tickborne diseases are increasing in some areas of the world, which has been linked to climate and land change (Gray et al., 2009;Madison-Antenucci et al., 2020). Our ability to reduce the likelihood of being infected in the field during restoration projects could be enhanced by employing drone-based preventative strategies. This sampling approach could be modified to survey other invertebrate species and communities as part of a restoration monitoring project.

| Air quality assessment
Similarly to using drones to sample soil and water samples, a drone can be mounted with an air sampling device to enable air quality assessments following a restoration intervention. Jumaah et al. (2021) developed a drone-based particulate matter at 2.5 μm (PM2.5) air quality monitoring system. The system included an air quality detector using Arduino sensors. The authors collected air PM2.5 data in a vertical flight path, which was similar to the ground truthed reference data. This could be useful for surveying large areas and collecting real-time air quality data following a restoration intervention (e.g., assessing whether a given restoration planting programme improves air quality). Several other researchers are developing drone-based air quality detection systems (Vijayakumar et al., 2020;Zhao et al., 2020). Hedworth et al. (2021) found that particle size distribution was unaffected by the drone and suggested that the area directly above the drone was optimal to mount an air quality sensor.
A similar method could collect biogenic compounds such as pollen and microbial communities to assess restoration outcomes on these biotic components of air. Indeed, collecting airborne eDNA samples is an emerging area of research in ecology, for example, for monitoring terrestrial vertebrate communities (Lynggaard et al., 2022).
Moreover, some researchers are assessing whether restoration interventions and aspects of habitat complexity can improve the aerobiome (microbiome of the air) with potential human health implications Robinson et al., 2020;Robinson et al., 2021). Drones could potentially help facilitate the collection of airborne eDNA samples in different restoration settings. Research is required to determine if this is better than traditional and less mobile methods (e.g., sampling stations with Petri dishes or handheld samplers).

| Digital models to assess post-restoration vegetation complexity
Following a restoration intervention (e.g., a restoration planting scheme), drones with LiDAR modules can help assess the intervention's success, as highlighted in Section 3.2. Along with LiDAR models, structure from motion could be useful in assessing vegetation biomass and complexity to aid restoration monitoring (Swinfield et al., 2019). As discussed in Section 3.2, combining drone hyperspectral and LiDAR data seems to be the most effective approach to assess forest structural attributes and tree species richness for restoration planning in a forest ecosystem. This approach can therefore F I G U R E 5 Modified drones can be used to collect samples in a range of environments-for example, (a) waterbodies, (b) soil, and (c) grasslands. contribute to several SER Recovery Wheel objectives (Figure 2), including Species Composition and Structural Diversity.

| SUMMARY OF P OTENTIAL CON S TR AINTS
It is important to re-emphasise the constraints associated with using drones in restoration. These include potential risks to wildlife, particularly if the drone operator is inexperienced and fails to follow operating guidelines for given species or communities, for example, for birds (Rümmler et al., 2018;Vas et al., 2015). The cost of the hardware is also a potential constraint, where drones (and modules/ sensors) themselves can be costly, particularly if damaged in the absence of insurance. Software, analysis and labour are also costly, plus methodological ambiguity can inhibit the generalisation of survey data (Buters, Bateman, et al., 2019). Public perception and opinion are also important factors; considering the social aspects of drone-assisted restoration, interventions will likely be critical to many projects' success and the continued development of dronebased methods in restoration. Research on the perception of drones in science in general has suggested that people can be apprehensive towards them, given their military origins (Markowitz et al., 2017). However, the same study showed moderate to strong public support for using drones for conservation purposes. This perception needs to be maintained-and potentially enhanced-if drones are to be used to help restore ecosystems in the long term. In many countries,

CO N FLI C T O F I NTE R E S T
The authors declare that the research was conducted without any commercial or financial relationships that could be construed as a potential conflict of interest.

PE E R R E V I E W
The peer review history for this article is available at https://publo ns.com/publo n/10.1111/2041-210X.13912.

DATA AVA I L A B I L I T Y S TAT E M E N T
There are no data associated with this manuscript.